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Used to represent the so called sonus medius, a short vowel sound between U and I before labial consonants in Latin words such as ⟨optͱmus⟩ for optumus and optimus, later used as a variant of y in inscriptions for Greek upsilon (as in ⟨Olͱmpicus⟩ for Olympicus), possibly inspired by an early form of the spiritus asper.
