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1977 May 1, Frank Wunderlich & Gerhard Herlan, “A reversibly contractile nuclear matrix”, in Journal of Cell Biology, volume 73, pages 271–278:From Tetrahymena macronuclei we have isolated a reversibly contractile nucleo-skeleton, i.e., an "expanded" nuclear matrix which reversibly contracts when the total concentration of the bivalent cations, Ca and Mg (3:2), is decreased to 5 mM or increased to 125 mM.
1998 April 24, Angus I. Lamond & William C. Earnshaw, “Structure and Function in the Nucleus”, in Science, volume 280, number 5363, →DOI, pages 547–553:At one extreme, the nucleus has been proposed to have its own nucleoskeleton and distinct organelles.
1999 June 11, Peter R. Cook, “The Organization of Replication and Transcription”, in Science, volume 284, number 5421, →DOI, pages 1790–1795:Finally, electron microscopy of chromatin-depleted nuclei shows that newly made DNA is initially associated with electron-dense bodies (diameters of 100 to 1000 nm) strung along a "nucleoskeleton"; with time, this DNA is extruded from these structures into adjacent regions (21 ).