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1992, Anna Maryańska-Nadachowska, Valentina G. Kuznetsova, Elżbieta Warchałowska-Śliwa, “Karyotypes of Psyllina (Homoptera)”, in Folia Biologica: International Journal of Biological Research, volume 40, Cracow Zoological Publications, page 23:
In all the species studied holokinetic chromosomes were found, this being a characteristic feature of the Hemiptera [...]