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Probably contracted from the genitive ending *-n + *ülä(“upper, above”) + Proto-Uralic*-ŋ(lative suffix), perhaps through a Uralic postposition *üleŋ: *-n-ülä-ŋ/*-n-üle-ŋ > *-n-leŋ > *-lle.[1]
In the past also thought to derive from *-la + the aforementioned lative suffix, in which case the long *-ll- would have been analogous to the adessive ending *-lla.