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1989, Kathleen Munthe, The Skeleton of the Borophaginae (Carnivora, Canidae): Morphology and Function, Publications in Geological Sciences, Volume 133, University of California Press, page 66,
Except in Borophagus and late Hemphillian Osteoborus, there is an entepicondylar foramen in borophagines, found in no living canid or hyaenid, but characteristic of felid humeri. In C. lupus and borophagines the lateral epicondyle bears two or more facets for the origin of the carpal and digital extensors and the supinator muscle.
1993, Frederick S. Szalay, Michael J. Novacek, Malcolm C. McKenna, Mammal Phylogeny: Placentals, Springer-Verlag, page 69:
The arctoids and early borophagine canids are the hallmark of the North American Mid-Miocene epoch.
2007, X. Wang, R. H. Tedford, “1: Evolutionary History of Canids”, in Per Jensen, editor, The Behavioural Biology of Dogs, CABI, page 10:
The history of the borophagines also begins with a small fox-like form, Archaeocyon, in the late Oligocene.