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No points were used by the ancient printers, excepting the colon and the period; but, after some time, a short oblique stroke, called a virgil, was introduced, which answered to the modern comma. In the fifteenth century this punctuation was improved by the famous Aldus Manutius with the typographical art in general; when he gave a better shape to the comma, added the semicolon, and assigned to the former points more proper places.
2004, Scott Shalaway, “Close-ups”, in Butterflies in the Backyard, Mechanicsburg, Pa.: Stackpole Books, →ISBN, page 18:
Commas (Polygonia comma) and Question Marks (Polygonia interrogationis) occur from the Gulf Coast to Canada and west to the Rockies. [...] Question Marks and Commas are handsome butterflies with burnt orange and black markings. [...] On the underside of each hind wing of the Comma is a small, distinctive silver hook that resembles a comma.
2013, Ann Simpson, Rob Simpson, “Butterflies and Moths”, in Nature Guide to Shenandoah National Park (Falcon Pocket Guide), Guilford, Conn., Helena, Mont.: Falcon Guides, Globe Pequot Press, →ISBN, page 91:
Other members of this genus that are frequently encountered in the park are the eastern comma (P. comma) and question mark (P. interrogationis).
(music) A difference in the calculation of nearly identical intervals by different ways.
(rhetoric) In Ancient Greekrhetoric, a short clause, something less than a colon, originally denoted by comma marks. In antiquity it was defined as a combination of words having no more than eight syllables in all. It was later applied to longer phrases, e.g. the Johannine comma.
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In the works of Cicero and Quintilian, the untransliterated Greek κόμμα(kómma) is used for comma in the grammatical sense of “a division…of a period smaller than a colon”.
“comma”, in Charlton T. Lewis and Charles Short (1879) A Latin Dictionary, Oxford: Clarendon Press
comma in Charles du Fresne du Cange’s Glossarium Mediæ et Infimæ Latinitatis (augmented edition with additions by D. P. Carpenterius, Adelungius and others, edited by Léopold Favre, 1883–1887)