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Uncertain. Compare κύλλα(kúlla, “puppy (hapax)”), σκύλιον(skúlion, “dogfish”), σκύλλα(skúlla, “shark”), as well as Old Armenianցուլ(cʻul, “bull”), Old Irishcuilén(“puppy”), Lithuaniankalė̃(“bitch, female dog”), and Russianклей(klej). A Proto-Indo-European*(s)kol-(“animal”) is possible, but problematic. Based on the phonetic issues with connecting the similar words mentioned above, Beekes tentatively takes the word as Pre-Greek;[1] it is possible that some, all, or none of the cognates mentioned above are taken from the same substrate continuum.