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Reconstructed by Ringe[1] from an alternating suffix *-nō- (singular) ~ *-na- (nonsingular), from Proto-Indo-European*-néh₂- ~ *-nh₂-, from n-infix presents to laryngeal-final roots that were reanalysed as a separate suffix (as in several other branches).
This suffix was preserved as a separate class only in Gothic. In Old High German, it mostly merged with the third weak class, while in the other descendants it merged with the second weak class.