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From Proto-Indo-European*mat-ḗn ~ *mat-nés, from *mat-(“insect, worm, maggot”), possibly borrowed from a substrate language. Compare Proto-Slavic*motyľь(“butterfly”), Sanskritमत्कुण(matkuṇa, “bug”). Possibly also related to Proto-Finnic*mato(“worm”) (akin to Proto-Samic*muocē), in which case it would be a borrowing in either direction (although seeing as the Finnic term has a convincing Indo-Iranian etymology, a borrowing from Finnic into Germanic appears more likely).