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The original heteroclitic alternation of r/n was preserved. The plural forms preserve the zero-grade forms of the suffix, and are like those of a neuter a-stem noun.
heteroclitic, irregularDeclension of *watōr (heteroclitic, irregular)
Ringe, Donald (2006) From Proto-Indo-European to Proto-Germanic (A Linguistic History of English; 1), Oxford: Oxford University Press, →ISBN, page 197