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from the accusative, analogically after other stem classes (u- and i-stems), probably to avoid homonymy with the neuter ending *-o.
A variant of the former view first proposed by Zaliznyak, Dybo and Nikolayev holds that IE *-os became *-ə in Slavic, which developed into *-ъ in mainstream dialects but into -е(-e) in Old Novgorodian. In the latter view, the Old Novgorodian ending can be considered original, with generalisation of the front variant from the soft declension.
Some authors also speculated that *-os could have had both outcomes depending on some other factors, with subsequent generalisation of one or the other in different grammatical categories.
Widely agreed to have been inherited from Proto-Indo-European*-om, itself composed from the thematic vowel *-o- and the accusative singular ending *-m. A small minority of researches instead argued for an extension of the u-stem ending.
Suffix
*-ъ
Accusative singular desinence for masculine o-stems
Proto-Indo-European*-om, the supposed athematic genitive plural alongside thematic *-ōm < *-o-om. A variant of this view holds that also thematic stems had *-om without the thematic vowel in Proto-Indo-European.
Proponents of the former view have sometimes connected the neocircumflex intonation in Slovene, the stem short vowel in Czech and the Serbo-Croatian desinence -ā with the alleged Proto-Slavic length.
Suffix
*-ъ
Genitive plural desinence for hard stems of all classes.