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Unknown. Perchance related to *marati(“to stain, to sully”). It might be a substrate word spread from Poland and only late, while comparisons with other Indo-European words are difficult and prone to chance correspondence. However it has terminally ousted the other Slavic word for madder, *broščь, by the end of the Early Modern Age.
*-asъ is the expected Balto-Slavic form but is found only in some Old Czech documents; -axъ is found everywhere else and is formed by analogy with other locative plurals in -xъ. ** The second form occurs in languages that contract early across /j/ (e.g. Czech), while the first form occurs in languages that do not (e.g. Russian).
Trubachyov, Oleg, editor (1990), “*marěna”, in Этимологический словарь славянских языков [Etymological dictionary of Slavic languages] (in Russian), numbers 17 (*lъžь – *matješьnъjь), Moscow: Nauka, →ISBN, page 210
Vasmer, Max (1964–1973) “марена”, in Oleg Trubachyov, transl., Этимологический словарь русского языка [Etymological Dictionary of the Russian Language] (in Russian), Moscow: Progress
