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Unknown. Synonym to today’s standard Lithuanianaukšlė̃ and Latvianaûsleja, also auslēja, auslejs. The formal discrepancy may be the effect of parallel borrowing from a substrate, after having shared the name with a similar fish, perhaps *klenь, *klěnь, but this looks like the same word.
*-asъ is the expected Balto-Slavic form but is found only in some Old Czech documents; -axъ is found everywhere else and is formed by analogy with other locative plurals in -xъ. ** The second form occurs in languages that contract early across /j/ (e.g. Czech), while the first form occurs in languages that do not (e.g. Russian).
Russian: укле́йка(ukléjka), укле́я́(ukléjá), sometimes still укле́й(ukléj) and see there for remoter forms. Generally, but sometimes, one did not write this fish-name with ѣ(jě) pre-1918.
Kolomijec, V. T. (1983) Происхождение общеславянских названий рыб [The Origin of the Common Slavic Names of Fish] (К IX Международному съезду славистов) (in Russian), Kyiv: Naukova Dumka, pages 89-93