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Explaining the final i-affection in Brittonic is difficult.
Matasović's suggestion to derive the Brittonic forms from *dolyā does not work, since *-yācannot cause i-affection.
An i-stem *dolis, although finding parallels in other Indo-European o-grade *-is nouns, is troubled by the fact that short *i cannot cause final i-affection of any vowel other than *e. The plural of an i-stem, if reconstructed as monophthongized *-eyes > *-īs, can yield the Brittonic forms but Schrijver rejects such a development existing (he thinks the i-stem plural instead yielded Welsh -ydd).
De Vaan reconstructs *dalī, assuming the zero-grade of the root.[2] The suffix can cause Brittonic i-affection, but this Celtic word family must have had the o-grade throughout given the derived singulative *dolinyā and its Goidelic reflexes in Old Irish duilne, Middle Irish duille etc. This poses a problem since Proto-Indo-European *-ih₂ does not naturally use the o-grade of a root.
^ De Vaan, Michiel (2008) “folium”, in Etymological Dictionary of Latin and the other Italic Languages (Leiden Indo-European Etymological Dictionary Series; 7), Leiden, Boston: Brill, →ISBN, page 230