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Unknown. The forms in *-r- are difficult to explain; they may be from the addition of *-raz with metathesis (Kroonen, Orel) or conflation with *hairaz (Orel). Kroonen speculatively connects it with Albaniankirrem(“to become hoarse”), for Proto-Indo-European*keys-, *kis-r-, but adds that the Germanic words may all derive from *hissōną(“to hiss”), ultimately of onomatopoeic origin. Compare *hwistlōną(“to whistle”) for a similar (but formally unrelated) onomatopoeic root.
