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From Proto-Indo-European*kʷékʷlom(“wheel”). The plural *hweulō is from the Proto-Indo-European collective *kʷekʷléh₂(“wheels”), and the neuter gender was likely back-formed from that.
The plural *hweulō descends from an Indo-European collective noun and shows the original Verner alternant *gw, which later became *w (represented by *u).
neuter a-stemDeclension of *hwehwlą (neuter a-stem)