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From an earlier *želdlo, formed as *želti(“to bite, to tingle”) + *-dlo, from Proto-Balto-Slavic*geldla, from Proto-Indo-European*gʷelH-dʰlo-m, from *gʷelH-(“to quell, to stroke”). The nasalization is secondary, result of disimilation *-ldl- > *-ndl- (a similar development occurred in *dętelъ(“woodpecker”), from an earlier *delb-telъ).
Vasmer, Max (1964–1973) “жало”, in Oleg Trubachyov, transl., Этимологический словарь русского языка (in Russian), Moscow: Progress
Georgiev, Vladimir I., editor (1971), “жело”, in Български етимологичен речник (in Bulgarian), volume 1 (А – З), Sofia: Bulgarian Academy of Sciences Pubg. House, →ISBN, page 533
Georgiev, Vladimir I., editor (1971), “жило”, in Български етимологичен речник (in Bulgarian), volume 1 (А – З), Sofia: Bulgarian Academy of Sciences Pubg. House, →ISBN, page 546
References
^ Derksen, Rick (2008) “žędlo”, in Etymological Dictionary of the Slavic Inherited Lexicon (Leiden Indo-European Etymological Dictionary Series; 4), Leiden, Boston: Brill, →ISBN, page 560