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The Psalterium Sinaiticum contains the only Old Church Slavonic instance of бьчела(bĭčela) but since the form бъчела(bŭčela) is just as rare, it is impossible to precisely reconstruct the late Proto-Slavic form of this word on the basis of Old Church Slavonic. Since this Proto-Slavic word was an oxytone, none of the descendant languages retain a reflex of the medial weakjer.
Alternatively (obsolete) explained as a deverbative of *bučati(“to make noise, roar”) and reconstructed as *bъčela. A bee is, therefore, the one that makes noise.
*-asъ is the expected Balto-Slavic form but is found only in some Old Czech documents; -axъ is found everywhere else and is formed by analogy with other locative plurals in -xъ. ** The second form occurs in languages that contract early across /j/ (e.g. Czech), while the first form occurs in languages that do not (e.g. Russian).