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(sciences, by extension) Inserted between other things
1982 December, I. F. Zhimulev et al., “Intercalary heterochromatin in Drosophila”, in Chromosoma, volume 87, →DOI, pages 197–228:
Breaks caused by chromosome aberrations in regions with repeats may not result in a sharp decline of viability, so that break points of chromosome rearrangements in intercalary heterochromatin may be more frequent than in other regions.
2007 February 28, Kitokazu Agata et al., “Unifying principles of regeneration I: Epimorphosis versus morphallaxis”, in Development, Growth, and Differentiation, →DOI:
In this new model, the anterior and posterior blastemas were formed as signaling centers to direct intercalary reorganization of body regionality or positional information.
(botany) of a meristem: situated between zones of permanenttissue, thus a shoot growing at the base of a leaf, in comparison with apical growth at the tip of a root or plant.
(entomology) of a wing vein: between the major veins common to insect wings.
1953, Willis W. Wirth, “American Biting Midges of the Helid Genus Monohelea”, in Proceedings of the United States National Museum, volume 103, number 3320, pages 135–154:
...two radial cells, second longer than first; intercalary fork fairly distinct; crossvein r-m vertical...