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(biology) A surface on which an organism grows, or to which an organism or an item is attached.
The rock surface of a rockpool is the substrate for a sessile organism such as a limpet.
2000, Mike Hansell, Bird Nests and Construction Behaviour, →ISBN, page 90:
This definition [of "tool"] is not simple, but contains several elements. The tool must not be part of the animal's body (a beak is not a tool); the user must manipulate the tool in some way for it to realise its function; and, finally, a tool cannot be attached to the substrate. This is a fairly clear definition, but does seem to produce some rather arbitrary distinctions (Hansell 1987b). The spider Dinopis, for example, makes a small web which it holds in its legs, thrusting it down on passing ants. This is a tool, but all other webs, however complex, are not since they are anchored to the substrate. The woodpecker finch […] that uses a fine stick held in the beak to extract insect prey from wood, is a tool user, but a shrike […] that impales an insect on a thorn still attached to the bush is not.
2006, Edward A. Wasserman, Thomas R. Zentall, Comparative Cognition: Experimental Explorations of Animal Intelligence, →ISBN, page 520:
Detach/subtract [tasks involve] Severing a fixed attachment between environmental objects (or the substrate) or removing object(s) from another unattached object, so the latter is a more useful tool.
2015 December 29, “Spatial Patterns in the Distribution, Diversity and Abundance of Benthic Foraminifera around Moorea (Society Archipelago, French Polynesia)”, in PLOS ONE, →DOI:
Amphisteginids and peneroplids were among the few taxa found in the bay environments, probably due to their preferences for phytal substrates and tolerance to moderate levels of eutrophication.
(linguistics) A language that is replaced in a population by another language and that influences the language imposed on its speakers.
(plating) A metal which is plated with another metal which has different physical properties.
(construction) A surface to which a substance adheres.
Synonyms
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1663, Robert Boyle, “(please specify the page)”, in Some Considerations Touching the Vsefulnesse of Experimental Naturall Philosophy,, Oxford, Oxfordshire: Hen Hall printer to the University, for Ric Davis, →OCLC, part I:
The melted glass being supported by the substrated sand.