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Uncertain. Various explanations have been proposed, none without problems.
From *ḱel-(“to cover”), the root of *helaną.[1] The regular outcome of the zero grade would be *hl-, but it is possible that u was inserted to avoid a nonsyllabic root, by analogy with many other Germanic zero grades.
From *ḱewh₁-(“to swell”).[2] A long vowel would be expected in the zero grade due to the following laryngeal, but Kroonen explains this away as "pretonic shortening". In this case, related to Latincavus and Ancient Greekκοῖλος(koîlos).
^ Philippa, Marlies, Debrabandere, Frans, Quak, Arend, Schoonheim, Tanneke, van der Sijs, Nicoline (2003–2009) “hol 1”, in Etymologisch woordenboek van het Nederlands (in Dutch), Amsterdam: Amsterdam University Press
^ Guus Kroonen (2013) “*hula-”, in Alexander Lubotsky, editor, Etymological Dictionary of Proto-Germanic (Leiden Indo-European Etymological Dictionary Series; 11), Leiden, Boston: Brill, →ISBN, page 253