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Of unclear origin. Likely a borrowing, along with नक्र(nakra, “Scorpio; alligator; sea monster”), to which it may be related, from a non-Indo-European source, perhaps Dravidian. Attempts to connect the word with fish names in Hellenic and Baltic, as well as with Old Church Slavonicмокръ(mokrŭ, “wet”), are unconvincing.
Mayrhofer, Manfred (1996) “MAḾH- - mákara-”, in Etymologisches Wörterbuch des Altindoarischen [Etymological Dictionary of Old Indo-Aryan] (in German), volume II, Heidelberg: Carl Winter Universitätsverlag, page 286
Mayrhofer, Manfred (1963) “makamakāyate— mákarah”, in Kurzgefasstes Etymologisches Wörterbuch des Altindischen [A Concise Etymological Sanskrit Dictionary] (in German), volume II, Heidelberg: Carl Winter Universitätsverlag, pages 539-540