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From two sources, both reflecting earlier *-ōjaną:
From Proto-Indo-European*-eh₂yéti, an innovated compound suffix from *-éh₂(eh₂-stem noun suffix) + *-yéti(verbal suffix). These were originally formed as denominative verbs from ō-stem nouns. When attached to thematic (a-stem) nouns, the thematic vowel was also retained, but the resulting compound suffix *-eyé- became part of the first weak class instead.
From Proto-Indo-European*-(e)h₂yéti, an innovated compound suffix from *-(e)h₂ti(factitive verb suffix) which was originally athematic, but later extended with the thematic present suffix *-yéti.