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Traditionally reconstructed with leading *h₁, but several Greek and Sanskrit compounds, if the analysis is correct, rule this out, for example Ancient Greek ἀκαρός(akarós), a variant of ἔγκαρος(énkaros, “brain”), or Sanskrit हेमन्(héman, “in winter”) < *ǵʰeymén.
Derived terms
*h₁en-dró-s(“egg, scrotum”)
Proto-Balto-Slavic:
>? Proto-Slavic: *ędro(“kernel”) (see there for further descendants)
Proto-Indo-Iranian: *Hāndrás(unexpected -ā-?)
Proto-Indo-Aryan: *Hāṇḍrás
>? Sanskrit: आण्ड(āṇḍa), अण्ड(aṇḍa, “egg, testicle”) (see there for further descendants)
Proto-Nuristani: (per Morgenstierne, who reconstructs *ãŕa-)
^ Schrijver, Peter C. H. (1991) The reflexes of the Proto-Indo-European laryngeals in Latin (Leiden studies in Indo-European; 2), Amsterdam, Atlanta: Rodopi, →ISBN, page 59: “*h₁en-”
^ Beekes, Robert S. P. (2011) Comparative Indo-European Linguistics: An Introduction, revised and corrected by Michiel de Vaan, 2nd edition, Amsterdam, Philadelphia: John Benjamins Publishing Company, page 247: “*h₁en”
^ Ringe, Donald (2006) From Proto-Indo-European to Proto-Germanic (A Linguistic History of English; 1), Oxford: Oxford University Press, →ISBN, page 71: “PIE *en”
↑ 5.05.15.25.35.45.5Dunkel, George E. (2014), “Lexikon ”, in Lexikon der indogermanischen Partikeln und Pronominalstämme [Lexicon of Indo-European Particles and Pronominal Stems] (Indogermanische Bibliothek. 2. Reihe: Wörterbücher) (in German), volume 2, Heidelberg: Universitätsverlag Winter GmbH Heidelberg, →ISBN, pages 221-236
^ Wodtko, Dagmar S.; Irslinger, Britta; Schneider, Carolin (2008) Nomina im indogermanischen Lexikon [Nouns in the Indo-European Lexicon] (in German), Heidelberg: Universitätsverlag Winter, page 165
↑ 7.07.1Nussbaum, Alan J. (1986) Head and Horn in Indo-European (Untersuchungen zur indogermanischen Sprach- und Kulturwissenschaft. NF / Studies in Indo-European Language and Culture. New Series; 2), Berlin, New York: Walter de Gruyter, →DOI, →ISBN, page 189
^ Derksen, Rick (2008), “*ę̄drò”, in Etymological Dictionary of the Slavic Inherited Lexicon (Leiden Indo-European Etymological Dictionary Series; 4), Leiden; Boston: Brill, →ISBN, →ISSN, page 157
^ Mayrhofer, Manfred (1992), “*āṇḍá-”, in Etymologisches Wörterbuch des Altindoarischen [Etymological Dictionary of Old Indo-Aryan] (in German), volume 1, Heidelberg: Carl Winter Universitätsverlag, page 162
^ Turner, Ralph Lilley (1969–1985), “āṇḍá”, in A Comparative Dictionary of the Indo-Aryan Languages, London: Oxford University Press
↑ 13.013.1Beekes, Robert S. P. (2010), “ἔν”, in Etymological Dictionary of Greek (Leiden Indo-European Etymological Dictionary Series; 10), volume I, with the assistance of Lucien van Beek, Leiden, Boston: Brill, →ISBN, page 419
^ Adams, Douglas Q. (2013), “ek*”, in A Dictionary of Tocharian B: Revised and Greatly Enlarged (Leiden Studies in Indo-European; 10), Amsterdam, New York: Rodopi, →ISBN, pages 78: “*h₁en-h₃kʷ-o-”
↑ 15.015.115.2De Vaan, Michiel (2008), “in”, in Etymological Dictionary of Latin and the other Italic Languages (Leiden Indo-European Etymological Dictionary Series; 7), Leiden, Boston: Brill, →ISBN, page 300
^ Zair, Nicholas (2012) The reflexes of the Proto-Indo-European laryngeals in Celtic, Leiden: Brill, →ISBN, page 261
^ Ziegler, Sabine (1994), Alfred Bammesberger and Günter Neumann, editors, Die Sprache der altirischen Ogam-Inschriften [The language of the Old Irish Ogham inscriptions] (Historische Sprachforschung; Ergänzungsheft 36) (in German), Göttingen: Vandenhoeck & Ruprecht, →ISBN, page 103
^ Beekes, Robert S. P. (2010), “ἐνῑπή”, in Etymological Dictionary of Greek (Leiden Indo-European Etymological Dictionary Series; 10), with the assistance of Lucien van Beek, Leiden, Boston: Brill, →ISBN
^ Wodtko, Dagmar S.; Irslinger, Britta; Schneider, Carolin (2008), “*h₃ekʷ-”, in Nomina im indogermanischen Lexikon [Nouns in the Indo-European Lexicon] (in German), Heidelberg: Universitätsverlag Winter, page 370
^ Frisk, Hjalmar (1960), “ἐνῶπα”, in Griechisches etymologisches Wörterbuch (in German), volume 1, Heidelberg: Carl Winter, pages 526-27
^ Beekes, Robert S. P. (2010), “ἐνῶπα”, in Etymological Dictionary of Greek (Leiden Indo-European Etymological Dictionary Series; 10), with the assistance of Lucien van Beek, Leiden, Boston: Brill, →ISBN
^ Beekes, Robert S. P. (2010), “εἰς”, in Etymological Dictionary of Greek (Leiden Indo-European Etymological Dictionary Series; 10), with the assistance of Lucien van Beek, Leiden, Boston: Brill, →ISBN
^ Beekes, Robert S. P. (2010), “ἐντός”, in Etymological Dictionary of Greek (Leiden Indo-European Etymological Dictionary Series; 10), with the assistance of Lucien van Beek, Leiden, Boston: Brill, →ISBN
^ Derksen, Rick (2008), “*ę̄trò”, in Etymological Dictionary of the Slavic Inherited Lexicon (Leiden Indo-European Etymological Dictionary Series; 4), Leiden; Boston: Brill, →ISBN, →ISSN, page 158
^ Kroonen, Guus (2013), “*nurþera-”, in Etymological Dictionary of Proto-Germanic (Leiden Indo-European Etymological Dictionary Series; 11), Leiden, Boston: Brill, →ISBN
^ Kroonen, Guus (2013), “*niþera-”, in Etymological Dictionary of Proto-Germanic (Leiden Indo-European Etymological Dictionary Series; 11), Leiden, Boston: Brill, →ISBN
^ Beekes, Robert S. P. (2010), “ἀκαρός”, in Etymological Dictionary of Greek (Leiden Indo-European Etymological Dictionary Series; 10), with the assistance of Lucien van Beek, Leiden, Boston: Brill, →ISBN, pages 49-50
^ Václav Blažek (2019), “Agamemnon”, in Georgios K. Giannakis, Christoforos Charalambakis, Franco Montanari and Antonios Rengakos, editors, Studies in Greek Lexicography (Trends in Classics – Supplementary Volumes; 72), De Gruyter, →DOI, →ISBN, §3.1.1, page 123: “ἄκαρος ‘brain’
<*H₁n̥-k̂r̥H₂o- vs. ἔγκαρος ‘brain’ [Lycophron
{320–280 BCE}, Alexandra 1104; Alcaeus Messenius {197 BCE}] <*H₁en-k̂r̥H₂o-”
^ Derksen, Rick (2008) Etymological Dictionary of the Slavic Inherited Lexicon (Leiden Indo-European Etymological Dictionary Series; 4), Leiden; Boston: Brill, →ISBN, →ISSN, page 387
^ Mayrhofer, Manfred (1992) Etymologisches Wörterbuch des Altindoarischen [Etymological Dictionary of Old Indo-Aryan] (in German), volume 1, Heidelberg: Carl Winter Universitätsverlag, pages 166-67
^ Derksen, Rick (2015) Etymological Dictionary of the Baltic Inherited Lexicon (Leiden Indo-European Etymological Dictionary Series; 13), Leiden, Boston: Brill, →ISBN, pages 196–197
^ Adams, Douglas Q. (2013), “e(n)-”, in A Dictionary of Tocharian B: Revised and Greatly Enlarged (Leiden Studies in Indo-European; 10), Amsterdam, New York: Rodopi, →ISBN, pages 87-88: “PIE *h₁en”