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Phono-semantic compound (形聲 / 形声, OC*suː): semantic 馬(“horse”) + phonetic 蚤(OC*ʔsuːʔ). Sense “grieved, sad” is a phonetic loan.
Etymology 1
to harass, to bother
Schuessler, 2007 suggests it is a voiced variant of 搜(sōu, “move”), p. 450.
grieved, sad
Of Austroasiatic origin. Schuessler, 2007 makes a connection to Old Khmer(sook, “sorrow, affliction, grief, to be sad, to grieve”), p. 451. The loss of OC coda is attributed to the long vowel.